Biosynthesis of carbzole

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With scheme Figure 2. Heterologousexpression and in vitro analysis of the cqsB genes. a) HPLC analysis

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of the standard of 1 (i)and the culture extracts from S. lividans TK23:pSE-cqsB (ii), S. lividans

TK23:pSEcqsBDcqsB4 (iii), and S. lividans TK23:pSE101 (iv;negative control). b) HPLC analysis of the in

vitro coupling reaction of the CqsB1, CqsB2, and CqsB3 reaction mixture (i)and the CqsB1 and CqsB3

reaction mixture (ii). c) Proposed biosynthetic pathway of 1.

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